It's not Nintendo nor Insomniac. Reporter Jamin Brophy-Warren declares that The Maw and Splosion Man development studio Twisted Pixel has Pixar's knack for creating memorable, unreal characters with which we can empathize. Argument here: The Pixar of Video Games [Slate]
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I would have guessed Valve. Then again, I think that TF2 might have a little too much to do with that decision.